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GTSG Publications

The Global Tree Specialist Group members are authoring numerous publications focusing on various aspects of tree conservation. These contributions are from scientific journals, magazines, books and other reports. Below is a selection of publications showcasing the wide range of contributions to the advancement of knowledge in the field of global tree conservation. 

• Acevedo-Mairena, H., Orozco, K. and Álvarez-Clare, S. 2024. Quercus insignis, Plan de Acción para Costa Rica 2025-2034. The Morton Arboretum. Lisle, IL. USA. ISBN 978-9930-00-123-3
• Baldini, R. M, A. L. MacVean, G. Cristofolini, T. Daniel, A. Managlia and M. Galloni. 2019. Synopsis and typification of the names published by Antonio Bertoloni in Florula Guatimalensis and in preceding publications. Phytotaxa 430 (3):199-213.
• Beech, E., Hills, R. and Rivers, M. 2022. GlobalTree Portal: visualizing the State of the World’s trees. Oryx, 56(3):332-332 https://doi.org/10.1017/S0030605322000254
• Beech, E., Rivers, M., Oldfield, S. and Smith, P.P., 2017. GlobalTreeSearch: The first complete global database of tree species and country distributions. Journal of Sustainable Forestry, 36(5), pp.454-489. https://doi.org/10.1080/10549811.2017.1310049
• Beech, E., Rivers, M., Rabarimanarivo, M., Ravololomanana, N.,Manjato, N., Lantoarisoa, F., Andriambololonera, S., Ramandimbisoa, B., Ralimanana, H., Rakotoarisoa, S., Razanajatovo , H., Razafiniary, V., Andriamanohera, A., Randrianasolo, V., Rakotonasolo, F., Rakotoarisoa, A., Randriamamonjy, N., Rajaovelona, L., Rakotomalala,N., Randriamboavonjy, T., Rajaonah, M., Rabehevitra, D., Ramarosandratana, A.V., Rakotoarinivo, M., B.H. Ravaomanalina and Jeannoda, V. (2021). Red List of Trees of Madagascar. BGCI. Richmond, UK
• BGCI. 2021. State of the World’s Trees. Richmond, UK, BGCI. 52 pp. https://www.bgci.org/resources/bgci-tools-and-resources/state-of-the-worlds-trees/
• Cannon, C.H., Dhyani, A., Jin, C. and Rivers, M. 2023. The Global Tree Assessment provides a multifaceted view on the future of tree diversity conservation. Plants, People, Planet. https://doi.org/10.1002/ppp3.10392
• Crowley, D., Barstow, M., Rivers, M. and Harvey-Brown, Y. (2020) The Red List of Acer: revised and extended. BGCI, Richmond, UK.
• Darbyshire, I., Timberlake, J., Osborne, J., Rokni, S., Matimele, H., Langa, C., Datizua, C., de Sousa, C., Alves, T., Massingue, A., Hadj-Hammou, J., Dhanda, S., Shah, T. and Wursten, B., 2019. The endemic plants of Mozambique: diversity and conservation status. PhytoKeys 136: 45–96. https://doi.org/10.3897/phytokeys.136.39020
• Davies, K., Starnes, T. and Rivers, M., 2024. Methodology for identifying the potential Alliance for Zero Extinction (AZE) tree species on a global scale. Conservation Science and Practice, 6(8), p.e13198.
• DRYFLOR*, 2016. Plant diversity patterns and their conservation implications in neotropical dry forests. Science 353(6306), pp.1383-1387. *Latin American and Caribbean Seasonally Dry Tropical Forest Floristic Network. http://doi.org/10.1126/science.aaf5080
• El Zein, H., Stephan, J. M., Khater, C., Al-Zein, M. S., and Bou Dagher Kharrat, M. 2022. Aligning terrestrial habitat typology of Lebanon with EUNIS habitat classification. https://doi.org/10.1127/phyto/2022/0390
• Erkens, R. H. J., Blanpain, L. M. P., Carrascosa Jara, I., Runge, K., Verspagen, N., Cosiaux, A., and Couvreur, T. L. P., 2023. Spatial distribution of Annonaceae across biomes and anthromes: Knowledge gaps in spatial and ecological data. Plants, People, Planet, 5(4), pp. 520-535. https://doi.org/10.1002/ppp3.10321
• Fernández, E., J. Timyan, C. D. Bacon and A. Henderson, 2023. Copernicia ekmanii in Haiti. Palms 67 (1): 5-12.
• Gallagher, R.V., Allen, S.P., Govaerts, R., Rivers, M.C., Allen, A.P., Keith, D.A., Merow, C., Maitner, B., Butt, N., Auld, T.D. and Enquist, B.J. 2023. Global shortfalls in threat assessments for endemic flora by country. Plants, People, Planet. https://doi.org/10.1002/ppp3.10369
• Hammoud, Y., and Stephan, J. 2022. Novelty predictors for shrub (and climbers) ecological niche modeling, based on their successional stage. Ecological Informatics, 71, 101771. https://doi.org/10.1016/j.ecoinf.2022.101771
• Hedges, S. B., W. W. B. Cohen, J. Timyan and Z. Yang, 2018. Haiti’s biodiversity threatened by nearly complete loss of primary forest. Proc Natl Acad Sci USA 115 (46): 11850-11855. https://doi.org/10.1073/pnas.1809753115
• Helmstetter, A. J., Ezedin, Z., de Lírio, E. J., de Oliveira, S. M., Chatrou, L. W., Erkens, R. H. J., Larridon, I., Leempoel, K., Maurin, O., Roy, S., Zuntini, A. R., Baker, W. J., Couvreur, T. L. P., Forest, F., and Sauquet, H. 2025. Toward a phylogenomic classification of magnoliids. American Journal of Botany, 112(1), pp. e16451. https://doi.org/10.1002/ajb2.16451
• Hepner, S., Agonvonon, G. A., Ehbrecht, M., Iheaturu, C., Azihou, A. F., and Ifejika Speranza, C. (2025). Degradation and Fragmentation Effects on Structural Complexity in West‐African Forest Patches. Biotropica, 57(2), e70012. https://doi.org/10.1111/btp.70012
• Hills, R., Marfleet, K., Rivers, M., Beech, E., Barstow, M., Davies, K., Crowley, D., de Kok, R. and Prance, G., 2023. Raising awareness through the Global Tree Assessment: Extinction risk of six fully assessed tree groups. Plants, People, Planet, 5(4), pp.563-573. https://doi.org/10.1002/ppp3.10289
• Hong, F., S. Blair Hedges, Z. Yang, J. W. Suh, J. Timyan, Z. Zhu, 2025. Decoding primary forest changes in Haiti and the Dominican Republic using Landsat time series. Remote Sensing of Environment 318 (2023). https://doi.org/10.1016/j.rse.2024.114590
• Iheaturu, C. J., Hepner, S., Batchelor, J. L., Agonvonon, G. A., Akinyemi, F. O., Wingate, V. R., Speranza, C. I. (2024). Integrating UAV LiDAR and multispectral data to assess forest status and map disturbance severity in a West African forest patch. Ecological Informatics, 102876. https://doi.org/10.1016/j.ecoinf.2024.102876
• Jestrow, B., M. Hass, W. Cinea, A. R. Franck, J. C. Timyan, M. P. Griffith and J. F. Ortega, 2023. The Critically Endangered Haitian endemic palm Attalea crassispatha (Arecaceae) and its living collections in Fairchild Tropical Botanic Garden: Perspectives from conservation surveys and DNA microsatellite (SSR) data. Webbia: J. Plant Tax. and Geogr. 78 (1): 3-14. https://doi.org/10.36253/jopt-14606
• Jose, J. K. 2024. India’s tree-planting strategies fall short. Science, 385(6711), 836-837. https://doi.org/10.1126/science.adq6755
• Jose, J. K. 2024. India’s Forest Conservation Amendment Act raises important questions. Frontiers in Ecology and the Environment, 22(4), e2741. https://doi.org/10.1002/fee.2741
• Jose, J. K. 2024. Persisting plant blindness in conservation efforts. Integrative Conservation. https://doi.org/10.1002/inc3.73
• Jose, J. K. 2025. The need for uprooting offensive plant eponyms. Botanical Journal of the Linnean Society, boaf008. https://doi.org/10.1093/botlinnean/boaf008
• Jose, J., Anuraj, K., and Gokul, K. 2023. Conservation of Diospyros crumenata (Ebenaceae), an Endangered tree endemic to the Western Ghats, India. Oryx, 57(4), 424-424. https://doi.org/10.1017/S0030605323000388
• Kress, W.J., Fér, T., Carlsen, M.M., and Krupnick, G.A. 2025. A conservation assessment and phylogenomic evaluation of protection status in the tropical genus Heliconia L. (Zingiberales: Heliconiaceae). Plants, People, Planet. https://doi.org/10.1002/ppp3.70000
• Kress, W.J. and Krupnick, G.A. 2022. Lords of the biosphere: Plant winners and losers in the Anthropocene. Plants, People, Planet, 4(4), pp. 350-366. https://doi.org/10.1002/ppp3.10252
• Krupnick, G.A. 2024. Gradations of Extinction. Arnoldia, 81(2), pp. 38–45. Veron, S., Bernard, A., Lebreton, E., Rodrigues-Vaz, C., Durand, M., Procopio, L., Helion, M., Gayot, M., Viscardi, G., Krupnick, G.A., Carrington, C.M.S., Boullet, V., Mallet, B., Dimassi, A., Pailler, T., Hivert, J., Lebouvier, M., Agnola, P., Bruy, D., Gateble, G., Lannuzel, G., Meyer, S., Gargominy, O., Gigot, G., Invernon, V. et al. 2023. Pre-assessments of plant conservation status in islands: the case of French Overseas Territories. Biodiversity and Conservation, 32, pp. 1165-1187. https://doi.org/10.1007/s10531-023-02544-8
• León, B., Roque, J., Ulloa Ulloa, C., Pitman, N., Jørgensen, P.M., and Cano, A. (eds.). 2006. Libro rojo de las plantas endémicas del Perú [Red book of Peru’s endemic plants]. Revista Peruana de Biología 13(2): 1-976.
• León-Yánez, S., Valencia, R., Pitman, N., Endara, L., Ulloa Ulloa, C., and Navarrete, H. (eds.). 2011. Libro rojo de las plantas endémicas del Ecuador, 2a edición. [Red book of Ecuador’s endemic plants, 2d edition.] Herbario QCA, Pontificia Universidad Católica del Ecuador, Quito.
• Linsky, J., Byrne, A., Handley, V., Coffey, E.D., Alvarez-Clare, S., Crowley, D., Meyer, A. (2024). Integrated plant conservation through the Global Conservation Consortia. Applications in Plant Sciences. e11586. https://doi.org/10.1002/aps3.11586
• Linsky, J. 2021. Prioritization to Action: How Red List Assessments are Guiding Coordinated Conservation Efforts in Magnolias. BGjournal 18(2): 35-37.
• Linsky, J., Coffey, E. E. D., Beech, E., Rivers, M., Cicuzza, D., Oldfield, S., and Crowley, D. 2022. Assessing Magnoliaceae through time: Major global efforts to track extinction risk status and ex situ conservation. Plants, People, Planet,1–6. https://doi.org/10.1002/ppp3.10285
• Linsky, J., Crowley, D., Beckman Bruns, E. and Coffey, E.E.D. 2022. Global Conservation Gap Analysis of Magnolia. Atlanta, GA: Atlanta Botanical Garden.
• MacDougal, J. and A. L. MacVean. 2022. The Mesoamerican Passiflora ilamo (Section Decaloba, Passifloraceae) newly recognized at species level. Novon 30:115-121. https://doi.org/10.3417/2022734
• MacVean A.L & H. Zinn. 2023. Plantas introducidas a Guatemala. In: Schuster J. C., J. Yoshimoto & J. Monzón Sierra (eds.) Biodiversidad de Guatemala, Vol 3. pp. 368-384. Universidad del Valle de Guatemala, Guatemala.
• Mapaura, A. and Timberlake, J.R., 2002. Zimbabwe. In: Southern Africa Plant Red Data Lists (edited J.S. Golding), pp. 157‒182. Southern African Botanical Diversity Network Report No. 14. SABONET, Pretoria.
• Marinho, L.C. and Beech, E., 2020. How phantom databases could contribute to conservation assessments. The Science of Nature, 107(3), p.21. https://doi.org/10.1007/s00114-020-01679-w
• Martén‐Rodríguez, S., Cristobal‐Pérez, E.J., De Santiago‐Hernández, M.H., Huerta‐Ramos, G., Clemente‐Martínez, L., Krupnick, G., Taylor, O., Lopezaraiza‐Mikel, M., Balvino‐Olvera, F.J., Sentíes‐Aguilar, E.M., Díaz‐Infante, S., Aguirre Jaimes, A., Novais, S., Cortés‐Flores, J., Lobo‐Segura, J., Fuchs, E.J., Delgado‐Carrillo, O., Ruiz‐Mercado, I., Sáyago‐Lorenzana, R., Pérez‐Arroyo, K., and Quesada, M. 2025. Untangling the complexity of climate change effects on plant reproductive traits and pollinators: A systematic global synthesis. Global Change Biology, 31(2). https://doi.org/10.1111/gcb.70081
• Mitri, G., Bechara, J., Stephan, J., Naimeh, A., McKenna M., Choueiter, D. 2023. Geoinformation applications for monitoring protected areas in Lebanon. Ninth International Conference on Remote Sensing and Geoinformation of the Environment (RSCy2023) 2023-09-21. https://doi.org/10.1117/12.2681075
• Moonlight, P.; K. Banda-R, O. Phillips, K. Dexter, R.T. Pennington, T. Baker, H. de Lima, L. Fajardo, R. González-M., R. Linares-Palomino, J. Lloyd, M. Nascimento, D. Prado, C. Quintana, R. Riina, G. Rodríguez M., D. Villela, A. Aquino, L. Arroyo, C. Bezerra, A. Brunello, R. Brienen, D. Cardoso, K.-J. Chao, Í.A. Cotta Coutinho, J. Cunha, T. Domingues, M. do Espírito Santo, T. Feldpausch, M. Ferreira Fernandes, Z. Goodwin, E. Jiménez, A. Levesley, L. Lopez-Toledo, B. Marimon, R. Miatto, M. Mizushima, A. Monteagudo, M. de Moura, A. Murakami, D. Neves, R. Nicora Chequín, T. de Sousa Oliveira, E. de Oliveira, L.P. de Queiroz, A. Pilon, D. Marques Ramos, C. Reynel, P. Rodrigues, R. Santos, T. Särkinen, V. da Silva, R. Souza, R. Vasquez, E. Veenendaal, 2021. Expanding tropical forest monitoring into Dry Forests: The DRYFLOR protocol for permanent plots. Plants, People, Planet, 3(3), pp.295-300. https://doi.org/10.1002/ppp3.10112
• Naznin, T. and A. L. MacVean. 2023. Impact of UV light on poinsettia (Euphorbia pulcherrima) architecture and quality. Acta Horticulturae 1377: 411-417.
• Özcan, A. U., Gülçin, D., Tuttu, G., Velázquez, J., Ayan, S., Stephan, J., Tuttu, U., Varlı, A., and Çiçek, K. (2024). The Future Possible Distribution of Kasnak Oak (Quercus vulcanica Boiss. and Heldr. ex Kotschy) in Anatolia under Climate Change Scenarios. Forests, 15(9), 1551. https://doi.org/10.3390/f15091551
• Pathak, H.N. and Bhuju, D.R., 2018. Secondary succession and soil nutrient dynamics in abandoned fallows of goth in Panchase, central midhill, Nepal. Journal of Natural History Museum, 30, pp.256-268.
• Pathak, H.N., 2014. Study of Myrsine capitellata wall in forest areas of Paiyunpata village, Baglung, Nepal. International Journal of Environment, 3(2), pp.55-62.
• Pathak, H.N., Bhuju, D.R., Shrestha, B.B. and Ranjitkar, S., 2021. Impacts of invasive alien plants on ecosystem services of Ramsar lake cluster in middle mountain Nepal. Global Ecology and Conservation, 27, p.e01597.
• Pathak, H.N., Shrestha, B.B., Bhuju, D.R. and Bhandari, P., 2020. Floristic Diversity in the Lake Cluster of Pokhara Valley, Central Nepal. Journal of Natural History Museum, 31(1), pp.1-32.
• Pathak, H.N., Shrestha, B.B., Bhuju, D.R. and Subedi, D.S., 2021. Spatial distribution of invasive alien plants in Pokhara valley, Nepal. Pak. J. Bot, 53(3), pp.1015-1024.
• Pennington, R.; D. Prado; C. Pendry, 2000. Neotropical seasonally dry forests and Quaternary vegetation changes. Journal of Biogeography 27, pp.261-273. https://doi.org/10.1046/j.1365-2699.2000.00397.x
• Quintana, I., Rivers, M. and Davies, K., 2024. Conservation Action Tracker: A tool to identify and monitor conservation actions for tree species. Applications in Plant Sciences, p.e11579. https://doi.org/10.1002/aps3.11579
• Rivers, M., Beech, E., Bazos, I., Bogunić, F., Buira, A., Caković, D., Carapeto, A., Carta, A., Cornier, B., Fenu, G. and Fernandes, F. et al., 2019. European Red List of Trees. International Union for Conservation of Nature and Natural Resources (IUCN).
• Rivers, M., Newton, A.C., Oldfield, S. and Global Tree Assessment Contributors 2022. Scientists’ warning to humanity on tree extinctions. Plants, People, Planet. https://doi.org/10.1002/ppp3.10314
• Rojas-Sandoval, J., L. Ferrufino, R. Flores, P. Galán, O. López, A. MacVean, D. Rodríguez and E. Chacón Madrigal. 2022. Flora introduced and naturalized in Central America. Biological Invasions 25(2):1007-1021
• Sarkinen, T., J.R. Iganci, R. Linares-Palomino, M. Simon and D. Prado, 2011. Forgotten forests- issues and prospects in biome mapping using Seasonally Dry Tropical Forests as a case study. BMC Ecology, 11, 27. http://www.biomedcentral.com/1472-6785/11/27
• Steed-Mundin, O., Crowley, D., Quintana, I., Wenham, J. (2024). Conservation Gap Analysis of Nothofagus. Wakehurst, Kew.
• Stephan, J., and Korban, M. (2025). Ecological niche modelling using MaxEnt for riparian species in a Mediterranean context. Ecological Indicators, 171, 113167. https://doi.org/10.1016/j.ecolind.2025.113167
• Steege, H.T., Pitman, N.C.A., and 118 others. 2013. Hyperdominance in the Amazonian tree flora. Science 342, 1243092. https://doi.org/10.1126/science.1243092
• Toledo‐Aceves T, Bonilla‐Moheno M, Sosa VJ, López‐Barrera F, Williams‐Linera G. 2022. Leaf functional traits predict shade tolerant tree performance in cloud forest restoration plantings. Journal of Applied Ecology 59: 2274-2289. https://doi.org/10.1111/1365-2664.14128
• Toledo-Aceves, T., Günter, S., Guariguata, M. R., García-Díaz,M, & Zhunusova, E. 2022. Financial Revenues from Timber Harvesting in Secondary Cloud Forests: A Case Study from Mexico. Forests 13, 1496. https://doi.org/10.3390/f13091496
• Toledo Aceves, T., Meave, J.A., González-Espinoza, M., Ramírez-Marcial, N. 2011. Tropical montane cloud forests: current threats and opportunities for their conservation and sustainable management in Mexico. Journal of Environmental Management 92: 974- 981. https://doi.org/10.1016/j.jenvman.2010.11.007
• Toledo-Aceves, T., Sosa, V., Vásquez-Reyes, V., & Sáenz-Romero, C. 2025. Assisted migration of cloud forest trees: Unearthing the effects of climatic transfer distance. Journal of Environmental Management, 377, 124591. https://doi.org/10.1016/j.jenvman.2025.124591
• Toledo-Garibaldi, M., Gallardo-Hernández, C., Ulian, T., & Toledo-Aceves, T. 2023. Urban forests support natural regeneration of cloud forest trees and shrubs, albeit with limited occurrence of late-successional species. Forest Ecology and Management, 546, 121327. https://doi.org/10.1016/j.foreco.2023.121327
• The Brazil Flora Group, Gomes-da-Silva, J., Filardi, F. L. R., Barbosa, M. R. V., Baumgratz, J. F. A., Bicudo, C. E. M., Cavalcanti, T. B., Coelho, M. A. N., Costa, A. F., Costa, D. P., Dalcin, E. C., Labiak, P., Lima, H. C., Lohmann, L. G., Maia, L. C., Mansano, V. F., Menezes, M., Morim, M. P., Moura, C. W. N.,…Zuntini, A. R., 2022. Brazilian Flora 2020: Leveraging the power of a collaborative scientific network. Taxon, 71(1), pp. 178-198. https://doi.org/10.1002/tax.12640
• Timberlake, J.R., Fagg, C. and Barnes, R.D. (1999). Field Guide to the Acacias of Zimbabwe. CBC Publishing, Harare, Zimbabwe. 160 pp.
• Timberlake, J.R., Goyder, D., Crawford, F., Burrows, J., Clarke, G.P., Luke, Q., Matimele, H., Müller, T., Pascal, O., de Sousa, C. and Alves, T., 2011. Coastal dry forests in Mozambique. Plant Ecology and Evolution 144(2): 126‒137.
• Timberlake, J.R., Nobanda, N. and Mapaure, I., 1993. Vegetation survey of the communal lands – north and west Zimbabwe. Kirkia 14(2): 171‒270.
• Timyan, J. C., A-I. Bonifassi and J-M Exantus, 2024. Census of the American Flamingo Phoenicopterus ruber (Phoenicopteriformes: Phoenicopteridae) in Haiti. Novitates Caribaea 24, (2024): 1-10. https://doi.org/10.33800/mc.vi24.353
• Verspagen, N., and Erkens, R. H. J., 2023. A method for making Red List assessments with herbarium data and distribution models for species-rich plant taxa: Lessons from the Neotropical genus Guatteria (Annonaceae). Plants, People, Planet, 5(4), pp. 536-546. https://doi.org/10.1002/ppp3.10309
• Zuntini, A. R., Carruthers, T., Maurin, O., Bailey, P. C., Leempoel, K., Brewer, G. E., Epitawalage, N., Françoso, E., Gallego-Paramo, B., McGinnie, C., Negrão, R., Roy, S. R., Simpson, L., Toledo Romero, E., Barber, V. M. A., Botigué, L., Clarkson, J. J., Cowan, R. S., Dodsworth, S.,…Baker, W. J., Phylogenomics and the rise of the angiosperms. Nature, 629(8013), pp. 843-850. https://doi.org/10.1038/s41586-024-07324-0